I saw this error today when receiving an email with an attachment. When I replayed the email again it worked fine.
[ERROR] 01/19/2021 13:27:48 PST Failed to save attachment attachment-file-name-goes-here.pdf 409 {'message': 'is at 608e3eafff3a06c30226d7623c04eeef72b1b814 but expected d8f53a0171e52ef86aac5eaf888d7efbb4048c9d', 'documentation_url': 'https://docs.github.com/rest/reference/repos#create-or-update-file-contents'}
I saw this error today when receiving an email with an attachment. When I replayed the email again it worked fine.