This issue was fixed by cloning the v2.10.0 GeoServer GeoPackage Extension (gs_geopkg) to the WorldWind GeoPackage Extension (worldwind_gs_geopkg) and modifying the following classes:
GeoPackageGetMapOutputFormat: added correct tile set matrix bounds to the meta data; leveraged the use of the "flipy" format option to invert the row ordering of the tile set.
GeoServer's GeoPackage export feature does not write out a GeoPackage with a tile matix conforming to the OGC GeoPackage Encoding Standard.