I changed the first Zenodo link to the URL that always resolves to the latest version as opposed to the DOI (also a link) given in the example, which is for the specific version mentioned in the example (4.20).
I made these changes into PR on a PR in case you want to more easily preview the changes in RStudio.
See also comments specific passages which I didn't attempt to change.
Further emendations, including
The form of the journal citation as emended essentially matches that given in the "How to cite" section of https://onlinelibrary.wiley.com/action/showCitFormats?doi=10.1111%2Fgcbb.12420 with only a few minor differences.
I changed the first Zenodo link to the URL that always resolves to the latest version as opposed to the DOI (also a link) given in the example, which is for the specific version mentioned in the example (4.20).
I made these changes into PR on a PR in case you want to more easily preview the changes in RStudio.
See also comments specific passages which I didn't attempt to change.