Extraposition. csubj attachment is wrong (head is "brushing", should be "took")
It was the Olmec who worshipped a rain god, a maize god, and the feathered serpent so important in the future pantheons of the Aztecs (who called him Quetzalcoatl) and the Maya (to whom he was Kukulkan).
Looks like an it-cleft to me, so acl:relcl should be advcl:relcl
From examining https://universal.grew.fr/?custom=6593a69b24161