NOTE: The BIN pin must not be shorted directly to VCC or VSS and any pullup resistor on the BIN pin must be connected only to VDD and not an external voltage rail. If an external thermistor is used for temperature input, the thermistor should be connected between this pin and VSS.
When a pack is used with the board, the BIN FET Q1 is turned on, shorting BIN to VSS
According to the BQ27426 datasheet:
When a pack is used with the board, the BIN FET Q1 is turned on, shorting BIN to VSS